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William L. McLean III Fellowship

The William L. McLean III Fellowship A fellowship for the study of environmental sciences and ornithology.

The Academy of Natural Sciences and the Department of Biodiversity, Earth and Environmental Science (BEES) is pleased to offer a fellowship in support of original graduate research in the fields of environmental science and/or ornithology. A faculty member within BEES must support and approve the application. The fellowship is for one term or up to an academic year, which may cover stipend, fees, tuition and research expenses (such as supplies and travel). Funds may be used for laboratory or field support as well as the final stages of a PhD dissertation.


Current BEES PhD students who have passed candidacy, or will have passed candidacy by the end of Fall Quarter in the upcoming academic year, are eligible for this fellowship.

The successful applicant will be required to serve as a Teaching Fellow for one quarter of the academic year; the quarter will be determined in advance at the discretion of the BEES department.

Required Materials:

  • Curriculum vitae
  • Statement of research interests (less than one page)
  • Two- to three-page description of the project and any progress to-date
  • Budget for one year of fellowship support (do not include stipend or tuition)
  • Letters of support from graduate adviser and one additional faculty member

Please email completed application packets as one PDF to


The application deadline is May 15th, 2019, with selection made by late June. Funds will be available at the start of the following academic year.


  • Danielle Moyer
  • Johannes Krause
  • Matthew Halley
  • Elizabeth McHone
  • Kaitlin Baudier
  • Abigail Elizabeth Dominy
  • A. Panagopoulou
  • Y. Hu
  • Tom Radzio

McLean Fellowship Publications

  • Baudier, K. M., & O’Donnell, S. (2016). Structure and thermal biology of subterranean army ant bivouacs in tropical montane forests. Insectes Sociaux, 63(3), 467-476. doi:10.1007/s00040-016-0490-2
  • Baudier, K. M., & O’Donnell, S. (2017). Weak links: How colonies counter the social costs of individual variation in thermal physiology. Current Opinion in Insect Science, 22, 85-91. doi:10.1016/j.cois.2017.06.004
  • Hu, Y., Holway, D. A., Łukasik, P., Chau, L., Kay, A. D., Lebrun, E. G., . . . Russell, J. A. (2017). By their own devices: Invasive Argentine ants have shifted diet without clear aid from symbiotic microbes. Molecular Ecology, 26(6), 1608-1630. doi:10.1111/mec.13991
  • Panagopoulou, A., Meletis, Z. A., Margaritoulis, D., & Spotila, J. R. (2017). Caught in the Same Net? Small-Scale Fishermen's Perceptions of Fisheries Interactions with Sea Turtles and Other Protected Species. Frontiers in Marine Science, 4. doi:10.3389/fmars.2017.00180
  • Radzio, T. A., Cox, J. A., Spotila, J. R., & O'connor, M. P. (2016). Aggression, Combat, and Apparent Burrow Competition in Hatchling and Juvenile Gopher Tortoises (Gopherus polyphemus). Chelonian Conservation and Biology, 15(2), 231-237. doi:10.2744/ccb-1181.1
  • Radzio, T. A., Cox, J. A., & O'Connor, M. P. (2017). Behavior and conspecific interactions of nesting gopher tortoises (gopherus polyphemus). Herpetological Conservation and Biology, 12(2), 373-383.
  • Radzio, T. A., Cox, J. A., & O'connor, M. P. (2017). Hatching Success and Other Reproductive Attributes of Gopher Tortoises in Southwest Georgia. Chelonian Conservation and Biology, 16(1), 103-105. doi:10.2744/ccb-1166.1

Dissertations (Award Year)

  • Baudier, K. M (2017). Microhabitat and elevational patterns in thermal tolerance and thermoregulation of Neotropical army ants (Formicidae: Dorylinae). PhD. Dissertation, Drexel University, BIO/BEES. (2015-2016)
  • Dominy, Abigal Elizabeth (2015) Modeling underwater visual ability and varied color expression in the diamondback terrapin (Malaclemys terrapin) in relation to potential mate preference by females. PhD. Dissertation, Drexel University, BEES. (2014-2015)
  • Panagopoulou, A. (2017) Sea turtles and small scale fisheries: Designing conservation policies for a marine area on Crete, Greece. PhD. Dissertation, Drexel University, BEES. (2014-2015)
  • Hu, Y. (2015) The role of symbiotic bacteria in the evolution of ant herbivory. PhD. Dissertation, Drexel University, BIO. (2014-2015)